That's up to you. A Tungsol in V1, with all the other's stock, should sound great. I have an RFT in V1 and V2 of my Herbert. The V1 tube I put in for the improved tone. The V2 RFT I put in because the Chinese tube got real noisy, and I had an extra. I do like the sound better with the V2 RFT, but all other's are stock. Since I love the sound now, I'll probably replace the others with Chinese when they go bad. Although I have a nice Phillips balanced 12AX7 that I'll probably put in the phase slot.
Anyway, I would just put one in V1, but you could try that and then put the others in V2 & v3, and see which you like better.