Really depends on the amp too, and what sort of preamp stage you've got driving into the power section. It's a yin-yang thing. It's really hard to say "all things being equal", because it's really amp and preamp specific.
I love EL34's for the grinding, chewy, middy, rich, full, looser (careful here folks, I use this term conservatively), more aggressive crunching tones. In to the higher gain ranks, it's good, it's harmonically rich and has some of that swirly edge to it.
I went with 6550's in my VH4S - the amp is my high gain beast...and it SCREAMS with the 6550's. It's thick, potent, hot, full of sizzle, and handles the compression fairly well...if that makes any sense. They like to be driven hard, and their response to clean to mean is very linear - whereas the EL34's tend to have different echelons of break up along the way.
Further note: Although "power valve saturation" is a highly overused term and concept around here, EL34's do sound nice (to my ears) when getting driven into the dirt than the 6550's do. But conversely, the 6550's take more of a beating and therefore don't saturate as easily, so it's kind of a moot point.
V.